Moving Toward an Unstable Equilibrium: Saddle Nodes in Population Systems

J. M. Cushing, Brian Dennis, Robert A. Desharnais, R. F. Costantino


Abstract

We identify an unstable equilibrium with a two-dimensional stable manifold and a one-dimensional unstable manifold in a three state variable (larva, pupa, adult) insect population growth model. The saddle node forecasts that the time series of some initial numbers of larvae, pupae, and adults are drawn closely to the unstable equilibrium before approaching the asymptotic stable attractor (a two-cycle) while the time series of other initial points are not. Using two quantitative indices, we examine time series from a Tribolium experiment for evidence of the predicted saddle node. We conclude that a saddle node accounts for the transient dynamics in these data and for the differences between the transient behaviour of different replicates of the same experiment.



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Cushing, J.M., Dennis, B., Desharnais, R.A., and Costantino, R.F. 1998. Moving toward an unstable equilibrium: saddle nodes in population systems. Journal of Animal Ecology 67: 298-306.

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This work was supported in part by grants DMS-9306271, DMS-9319073, DMS-9625576, DMS-9616205, DMS-9981374, DMS-9973126, DMS-9981458, and DMS-9981423 from the U.S. National Science Foundation. All opinions expressed are those of the authors and not necessarily those of the NSF.

  Copyright © 1997-2002, Robert A Desharnais
Department of Biological Sciences
California State University, Los Angeles, CA, 90032-8201
Email: rdeshar@calstatela.edu